The game carries on the style of gameplay from Rayman Origins in which up to four players simultaneously make their way through various levels. Lums can be collected by touching them, defeating enemies, or freeing captured Teensies. Along with Rayman, Globox, and the Teensies returning as playable characters, players can now control a new female character named Barbara.
In addition to the main playable characters, Murfy the fairy, who first appeared in Rayman 2: The Great Escape, appears as an assist character. Murfy can perform various actions such as cutting through ropes, activating mechanisms, grabbing hold of enemies and assisting in gathering Lums. These offer a range of levels in which co-operation is required to progress, such as levels that mix platforming for Rayman and friends with rhythm gameplay for Murfy. In the PlayStation Vita and Wii U versions of the game, an additional player can control Murfy directly with touch controls, using either the Wii U GamePad or the Vita's touch screen. In the PlayStation 3 and Xbox 360 versions of the game, Murphy is controlled by the computer and can be prompted to interact with certain objects with button controls. Clearing a level once unlocks an 'Invaded' version, where the level is remixed with additional enemies or new objectives. The game will have over 120 levels to clear. The game will also feature 40 remastered levels from the original Rayman Origins.
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